At noon on a cloudless on June
21st the sun delivers about 100,000 lumens of
visible light. Simultaneously, the sun delivers an equal
amount of energy as heat. The bush tries to use the light
while staying cool.
Leaflets in the sun lose heat by
air cooling and evaporation. According to Lambers, the speed
of air cooling increases 4xs when the biggest part of a
leaflet halves. In this picture, the smallest leaflet air
cools 15/4 x 15/4 = 14 times faster than the largest leaflet.
In order to document the effects
of light intensity further, the most typical one of ten
representative end leaflets were chosen in each of five light
conditions. From left to right those conditions were: deep
shade, medium shade, light shade, partial sun, and full sun.
Nothing grows in deep shade,
except a little poison-oak. The poison-oak leaflets are held
horizontally, and do not overlap. The leaflets are paler, and
the veins less pronounced than in other situations. Deep shade
can be seen under the clusters of big trees in the center of
Los Osos Oaks Reserve. Deep shade illumination is about 1,000
lumens.
Poison-oak grows most vigorously
in light shade, at 20,000 lumens. The stems grow 5 feet a
year, toward the light; and the leaves are more apt to have a
waxy coat. However, the plants fruit best in partial sunlight.
Sun and shade leaves are common
in nature. Here is coast live oak: